Long ago, dragons were hunted to near extinction by the Order of St. George, a legendary society of dragon slayers. Hiding in human form and growing their numbers in secret, the dragons of Talon have become strong and cunning, and they're positioned to take over the world with humans none the wiser.

Ember stands with Riley and his band of rogue dragons, instead of becoming an assassin for Talon. Meanwhile, she's lost contact with her twin brother Dante, as well as with Garret, a former-enemy soldier who challenged her beliefs about her human side.