As Ayla studies to become a spiritual leader to Jondalar's people, the Zelandonii, she faces many challenges, including being separated from Jondalar and their young daughter, Jonayla, and experiences a revelation.

Jondalar and Ayla are two Ice Age travelers on an uncertain journey. The journey will require them to brave flash floods and glaciers, hungry hyenas and vengeful hunter women, and midnight visions and painful echoes from the past.